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This weekend I upgraded a Jira instance from one LTS release to the next (8.13.1 to 8.20.1).

Everything appeared to go well, but shortly afterwards users started complaining: Jira is not emailing certain users.

I checked, and sure enough: there's a situation where Jira 8.13.x and all prior releases would email, but Jira 8.20.x will not.

Specifically, this affects users like this one, who are not in a group like jira-users blessed with 'Application access':

WEB > 2021/11/01 > Jira 8.20.x gotcha: lost notifications > image2021-11-1_23-28-0.png

What should happen if a new issue is created, assigned to 'helpdesk', or with 'helpdesk' as a watcher?

Well, in Jira 8.13.x, a notification would go out to helpdesk@mycompany.com
In Jira 8.20.x no notification goes out.

The 8.20.x behaviour is actually correct. An account without Browse permission shouldn't get emails. Yet this important bug fix is not listed in the release notes, nor in the [174 bugs fixed between 8.13.1 and 8.20.1|]